Eric R. Pianka's Publications


Papers and Chapters by Eric R. Pianka

(Abstracts and Book Reviews excluded)



Link to Google Scholar Citations Summary

List of ERP's Books

List of ERP's Papers in Copeia

List of ERP's Papers in Ecology

List of ERP's Papers in The American Naturalist

List of ERP's Papers in The Western Australian Naturalist

Pianka, E. R. and H. M. Smith. 1959. Distributional records for certain Mexican and Guatemalan reptiles. Herpetologica 15: 119-120. Download pdf

Willson, M. F. and E. R. Pianka. 1963. Sexual selection, sex ratio, and mating system. American Naturalist 97: 405-407. Download pdf.

Pianka, E. R. 1966. Latitudinal gradients in species diversity: A review of concepts. American Naturalist 100: 33-46. (Reprinted in 1981 in C. F. Jordan (ed.) Tropical Ecology, Benchmark Papers in Ecology. Hutchinson Ross Publishing Company.) Download pdf

Pianka, E. R. 1966. Convexity, desert lizards, and spatial heterogeneity. Ecology 47: 1055-1059. Download pdf

MacArthur, R. H. and E. R. Pianka. 1966. On optimal use of a patchy environment. American Naturalist 100: 603-609. Selected as "This Week's Citation Classic" in Current Contents (Social and Behavioral Sciences) (1988), volume 20 (number 31): page 16, in Current Contents (Arts & Humanities) (1988), volume 10 (number 31): page 16, and in Current Contents (Agriculture, Biology & Environmental Sciences) (1988), volume 19 (number 31): page 16.) (Reprinted in 1991 in L. A. Real and J. H. Brown (eds.) Foundations of Ecology. Classic Papers with Commentaries. Univ. Chicago Press.) Download pdf of paper, Download Figure 1

Pianka, E. R. 1967. On lizard species diversity: North American flatland deserts. Ecology 48: 333-351. Download pdf

Pianka, E. R. 1968. Notes on the biology of Varanus eremius. Western Australian Naturalist 11: 39-44. Download pdf

Pianka, E. R. 1969. Notes on the biology of Varanus caudolineatus and Varanus gilleni. Western Australian Naturalist 11: 76- 82. Download pdf

Pianka, E. R. 1969. Habitat specificity, speciation, and species density in Australian desert lizards. Ecology 50: 498-502. Download pdf

Pianka, E. R. 1969. Sympatry of desert lizards ( Ctenotus) in Western Australia. Ecology 50: 1012- 1030. Download pdf

Pianka, E. R. and H. D. Pianka. 1970. The ecology of Moloch horridus (Lacertilia: Agamidae) in Western Australia. Copeia 1970: 90-103. Download pdf

<-- World's smallest monitor lizard, actual size, a neonate Varanus brevicauda (SVL = 52 mm, weight 1.4 gms).


Pianka, E. R. 1970. Notes on Varanus brevicauda. Western Australian Naturalist 11: 113-116. Download pdf


Pianka, E. R. 1970. Notes on the biology of Varanus gouldii flavirufus. Western Australian Naturalist 11: 141-144. Download pdf

Pianka, H. D. and E. R. Pianka. 1970. Bird censuses from desert localities in Western Australia. Emu 70: 17-22. Download pdf

Pianka, E. R. 1970. Comparative autecology of the lizard Cnemidophorus tigris in different parts of its geographic range. Ecology 51: 703-720. Download pdf

Pianka, E. R. 1970. On r and K selection. American Naturalist 104: 592-597. (Reprinted in 1978 in T. H. Clutton-Brock and P. H. Harvey (eds.) Readings in Sociobiology. Freeman. Selected as "This Week's Citation Classic" in Current Contents (1979), volume 10 (number 47): page 10.) Download pdf

Pianka, E. R. 1971. Comparative ecology of two lizards. Copeia 1971: 129-138. Download pdf

Pianka, E. R. 1971. Species diversity. Chapter in Topics in the Study of Life: The Bio Source Book, pp. 401-406. Harper and Row, New York. Download pdf



ERP implants a transmitter into the coelom of a Varanus tristis.

Pianka, E. R. 1971. Notes on the biology of Varanus tristis. Western Australian Naturalist 11: 180-183. Download pdf

Pianka, E. R. 1971. Ecology of the agamid lizard Amphibolurus isolepis in Western Australia. Copeia 1971: 527-536. Download pdf

Pianka, E. R. 1971. Notes on the biology of Amphibolurus cristatus and Amphibolurus scutulatus. Western Australian Naturalist 12: 36-41. Download pdf

Pianka, E. R. and R. B. Huey. 1971. Bird species density in the Kalahari and the Australian deserts. Koedoe 14: 123-130. Download pdf

Pianka, E. R. 1971. Lizard species density in the Kalahari desert. Ecology 52: 1024-1029. Download pdf

Pianka, E. R. 1972. Zoogeography and speciation of Australian desert lizards: An ecological perspective. Copeia 1972: 127-145. Download pdf

Pianka, E. R. and W. S. Parker . 1972. Ecology of the iguanid lizard Callisaurus draconoides. Copeia 1972: 493-508. Download pdf

Pianka, E. R. 1972. r and K selection or b and d selection? American Naturalist 106: 581-588. Download pdf

Parker, W. S. and E. R. Pianka. 1973. Notes on the ecology of the iguanid lizard, Sceloporus magister. Herpetologica 29: 143-152. Download pdf

Pianka, E. R. 1973. The structure of lizard communities. Annual Review of Ecology and Systematics 4: 53-74. Selected as "This Week's Citation Classic" in Current Contents (Agriculture, Biology & Environmental Sciences) (1988), volume 19 (number 35): page 18.) Download pdf

Parker, W. S. and E. R. Pianka. 1974. Further ecological observations on the western banded gecko, Coleonyx variegatus. Copeia 1974: 528-531. Download pdf

Huey, R. B., E. R. Pianka, M. E. Egan, and L. W. Coons. 1974. Ecological shifts in sympatry: Kalahari fossorial lizards (Typhlosaurus). Ecology 55: 304-316. Download pdf



Pickling assembly line in the Kalahari, 1970.
Left to right: Larry Coons, Ray Huey, and Eric Pianka.


Pianka, E. R. 1974. Niche overlap and diffuse competition. Proc. Nat. Acad Sci. 71: 2141-2145. (Reprinted in 1975 in R. H. Whittaker and S. A. Levin (eds.) Niche: theory and application, Benchmark Papers in Ecology. Dowden, Hutchinson and Ross.) Download pdf

Huey, R. B. and E. R. Pianka. 1974. Ecological character displacement in a lizard. American Zoologist 14: 1127-1136. Download pdf

Pianka, E. R. and W. S. Parker . 1975. Ecology of horned lizards : A review with special reference to Phrynosoma platyrhinos. Copeia 1975: 141-162. Download pdf

Wake, D. B., R. G. Zwiefel, H. Desauer, G. Nace, E. R. Pianka, G. B. Rabb, R. Ruibal, J. Wright, and G. R. Zug. 1975. Report of the Committee on Resources in Herpetology. Copeia 1975: 391-404.


Wake, D. B., R. G. Zwiefel, H. Desauer, G. Nace, E. R. Pianka, G. B. Rabb, R. Ruibal, J. Wright, and G. R. Zug. 1975. Collections of preserved amphibians and reptiles in the United States. Herpetological Circ. 3: 1-22.

Wake, D. B., R. G. Zwiefel, H. Desauer, G. Nace, E. R. Pianka, G. B. Rabb, R. Ruibal, J. Wright, and G. R. Zug. 1975. Recommendations for the management of herpetological museum collections. Herpetological Review 6: 34-36.

Pianka, E. R. 1975. Niche relations of desert lizards. Chapter 12 (pp. 292-314) in M. Cody and J. Diamond (eds.) Ecology and Evolution of Communities. Harvard University Press. Download pdf

Pianka, E. R. and W. S. Parker . 1975. Age-specific reproductive tactics. American Naturalist 109: 453-464. Download pdf

Parker, W. S. and E. R. Pianka. 1975. Comparative ecology of populations of the lizard Uta stansburiana. Copeia 1975: 615- 632. Download pdf

Pianka, E. R. and H. D. Pianka. 1976. Comparative ecology of twelve species of nocturnal lizards (Gekkonidae) in the Western Australian desert. Copeia 1976: 125-142. Download pdf

Parker, W. S. and E. R. Pianka. 1976. Ecological observations on the leopard lizard Crotaphytus wislizeni in different parts of its range. Herpetologica 32: 95-114. Download pdf

Pianka, E. R. 1976. Competition and niche theory. Chapter 7 (pp. 114- 141) in R. M. May (ed.) Theoretical Ecology: Principles and Applications. Blackwell.

Pianka, E. R. 1976. Natural selection of optimal reproductive tactics. American Zoologist 16: 775-784. Download pdf.

Huey, R. B. and E. R. Pianka. 1977. Natural selection for juvenile lizards mimicking noxious beetles. Science 195: 201-203. Download pdf

Pianka, E. R. 1977. Reptilian species diversity. Chapter 1 (pp. 1-34) in C. Gans and D.W. Tinkle (eds.) Biology of the Reptilia, Academic Press, New York. Download pdf

Huey, R. B. and E. R. Pianka. 1977. Patterns of niche overlap among broadly sympatric versus narrowly sympatric Kalahari lizards (Scincidae: Mabuya). Ecology 58: 119-128. Download pdf.

Huey, R. B. and E. R. Pianka. 1977. Seasonal variation in thermoregulatory behavior and body temperature of diurnal Kalahari lizards. Ecology 58: 1066-1075. (With an Appendix by J. A. Hoffman.) Download pdf.

Schall, J. J. and E. R. Pianka. 1977. Species densities of reptiles and amphibians on the Iberian peninsula. Donana, Acta Vertebrata 4: 27-34. Download pdf

Pianka, E. R. and R. B. Huey. 1978. Comparative ecology, niche segregation, and resource utilization among gekkonid lizards in the southern Kalahari. Copeia 1978: 691- 701. Download pdf

Schall, J. J. and E. R. Pianka. 1978. Geographical trends in numbers of species. Science 201: 679-686. Download pdf

Schoener, T. W., R. B. Huey, and E. R. Pianka. 1979. A biogeographic extension of the compression hypothesis: Competitors in narrow sympatry. American Naturalist 113: 295-298. Download pdf

Pianka, E. R. 1979. Diversity and niche structure in desert communities. Chapter 10 (pp. 321-341) in R. Perry and D.Goodall (eds.) Arid-Land Ecosystems: Structure, Function and Management, Cambridge University Press. Download pdf

Pianka, E. R., R. B. Huey and L. R. Lawlor. 1979. Niche segregation in desert lizards. Chapter 4 (pp. 67-115) in D. J. Horn, R. Mitchell, and G. R. Stairs (eds.) Analysis of Ecological Systems. Ohio State University Press, Columbus. Download pdf

Schall, J. J. and E. R. Pianka. 1980. Evolution of escape behavior diversity. American Naturalist 115: 551-566. Download pdf

Pianka, E. R. 1980. Konkurrenz und Theorie der okologischen Nische. Chapter 7 (pp. 105-128) in R. M. May (ed.) Theoretische Okologie. Verlag Chemie, Basel.

Pianka, E. R. 1980. Guild structure in desert lizards. Oikos 35: 194- 201. Download pdf

Huey, R. B. and E. R. Pianka. 1981. Ecological consequences of foraging mode. Ecology 62: 991-999. Download pdf

Pianka, E. R. 1981. Competition and niche theory. Chapter 8 (pp. 167- 196) in R. M. May (ed.) Theoretical Ecology, Second Edition, Blackwell. Download pdf

Pianka, E. R. 1981. Diversity and adaptive radiations of Australian desert lizards. Chapter 50 (pp. 1375-1392) in A. Keast (ed.) Ecological Biogeography in Australia. D. W. Junk, The Hague, Netherlands. (Reprinted in 1984 as Chapter 4.6 (pp. 371-376) in M. Archer and G. Clayton (eds.) Vertebrate Zoogeography and Evolution in Australasia. University of New South Wales, Kensington, N. S. W., Australia.) Download pdf

Pianka, E. R. and J. J. Schall. 1981. Species densities of terrestrial vertebrates in Australia. Chapter 59 (pp. 1675-1694) in A. Keast (ed.) Ecological Biogeography in Australia. D. W. Junk, The Hague, Netherlands. (Reprinted in 1984 as Chapter 1.11 (pp. 119-124) in M. Archer and G. Clayton (eds.) Vertebrate Zoogeography and Evolution in Australasia. University of New South Wales, Kensington, N. S. W., Australia.) Download pdf

Pianka, E. R. 1981. Resource acquisition and allocation among animals. Chapter 12 (pp. 300-314) in C. Townsend and P. Calow (eds.) Physiological Ecology: An Evolutionary Approach to Resource Use. Blackwell. Download pdf

Ricklefs, R. E., D. Cochran, and E. R. Pianka. 1981. A morphological analysis of the structure of communities of lizards in desert habitats. Ecology 62: 1474-1483. Download pdf

Pianka, E. R. 1982. Observations on the ecology of Varanus in the Great Victoria desert. Western Australian Naturalist 15: 37-44. Download pdf

Pianka, E. R. and W. F. Giles. 1982. Notes on the biology of two species of nocturnal skinks, Egernia inornata and Egernia striata, in the Great Victoria desert. Western Australian Naturalist 15: 44-49. Download pdf

Huey, R. B., E. R. Pianka, and T. W. Schoener. 1983. Introduction (pp. 1-6) in R. B. Huey, E. R. Pianka, and T. W. Schoener.(eds.) Lizard Ecology: Studies of a Model Organism. Harvard University Press.

Huey, R. B. and E. R. Pianka. 1983. Temporal separation of activity and interspecific dietary overlap (with an Appendix by S. L. Pimm). Chapter 13 (pp. 281-296) in Huey, R. B. , E. R. Pianka, and T. W. Schoener (eds.) Lizard Ecology: Studies of a Model Organism. Harvard University Press.

Huey, R. B., E. R. Pianka, and C. M. Cavalier. 1983. Ecology of Lizards in the Kalahari Desert, Africa. National Geographic Society Research Reports 16: 365- 370. Download pdf

Pianka, E. R. 1985. Ecological dynamics of Australian desert lizards: or, the L-area revisited. National Geographic Society Research Reports 18: 581-591. Download pdf

Pianka, E. R. 1985. Some intercontinental comparisons of desert lizards. National Geographic Research 1: 490-504. Download pdf

Pianka, E. R. 1985. A wild analogy. BioScience 35: 685. Download pdf

Pianka, E. R. 1986. Ecological phenomena in evolutionary perspective. Chapter 16 (pp. 325-336) in N. Polunin (ed.) Ecosystem Theory and Application. Wiley and Sons. Download pdf

Pianka, E. R. 1987. Waiting for the apple to fall, or pooling our brainpower. Bull. Ecol. Soc. America 68: 12-13. Download pdf

Pianka, E. R. 1987. The subtlety, complexity and importance of population interactions when more than two species are involved. Revista Chilena de Historia Natural 60: 351-362. Download pdf

Pianka, E. R. 1988. Destiny. Bull. Ecol. Soc. America 69: 33. Download pdf

Pianka, E. R. 1989. Latitudinal gradients in species diversity. Trends in Ecology and Evolution 4: 223.

Pianka, E. R. 1989. Desert lizard diversity: additional comments and some data. American Naturalist 134: 344-364. Download pdf.

Pianka, E. R. 1989. The role of plants in evolutionary ecology. Evolutionary Trends in Plants 3: 75-80. Download pdf

Winemiller, K. O. and E. R. Pianka. 1990. Organization in natural assemblages of desert lizards and tropical fishes. Ecological Monographs 60: 27-55. Download pdf.

Duellman, W. E. and E. R. Pianka. 1990. Biogeography of nocturnal insectivores: historical events and ecological filters. Annual Review of Ecology and Systematics 21: 57-68. Download pdf.

Pianka, E. R. 1991. Phrynosoma platyrhinos. Catalogue of American Amphibians and Reptiles. pp. 517.1-517.4.

Pianka, E. R. 1992. A land of lizards. Landscope 7 (3): 10-16. Download pdf.

Pianka, E. R. 1992. Fire Ecology. Disturbance, spatial heterogeneity, and biotic diversity: fire succession in arid Australia. Research and Exploration 8: 352-371.

Pianka, E. R. 1992. Evolution and the ecosystem. Pages 9-19 in S. Baba, O. Akerele, and Y. Kawaguchi (eds.) Natural Resources and Human Health - Plants of medicinal and nutritional value. Proceedings of the First World Health Organization Symposium on Plants and Health for all: Scientific Advancement, Kobe, Japan, 26-28 August 1991. Elsevier Science Publishers.

Pianka, E. R. 1992. The state of the art in community ecology. In K. Adler (ed.) Herpetology. Current Research on the Biology of Amphibians and Reptiles. Proceedings of the First World Congress of Herpetology at Canterbury. Contributions to Herpetology, Number 9: 141-162. Download pdf.

Pianka, E. R. 1992. Reproductive tactics. Pages 189-209 in R. Dallai (ed.) Sex origin and evolution. Proc. of International Symposium on Origin and Evolution of Sex, Siena, September 9-11, 1991. Mucchi Editore, Italy. Download pdf.

Pianka, E. R. 1993. The many dimensions of a lizard's ecological niche. Chapter 9 (pp. 121-154) in E. D. Valakos, W. Bohme, V. Perez-Mellado, and P. Maragou (eds.) Lacertids of the Mediterranean Basin. Hellenic Zoological Society. University of Athens, Greece. Download pdf

Haydon, D. T. , R. R. Radtkey, and E. R. Pianka. 1993. Experimental Biogeography: Interactions between stochastic, historical, and ecological processes in a model archipelago. Chapter 11 (pp. 117-130) in R. E. Ricklefs and D. Schluter (eds.) Species Diversity in Ecological Communities: Historical and Geographical Perspectives. University of Chicago Press. Download pdf

Heatwole, H. and E. R. Pianka. 1993. Natural History of the Squamata. Pages 197-209 in G. J. B. Ross (ed.) Fauna of Australia. Volume 2. Amphibia Reptilia Aves. Australian Biological and Environmental Survey, Canberra. Download pdf

Vitt, L. J. and E. R. Pianka. 1994. Introduction and acknowledgments. Pages ix-xii in Vitt, L. J. and E. R. Pianka (eds.), Lizard Ecology: Historical and Experimental Perspectives. Princeton University Press. 403 pp.

Haydon, D. T., B. I. Crother, and E. R. Pianka. 1994. New directions in biogeography? Trends in Ecology and Evolution 9: 403-406. Download pdf.

Pianka, E. R. 1994. Comparative ecology of Varanus in the Great Victoria desert. Australian Journal of Ecology 19: 395-408. Download pdf.

Pianka, E. R. 1994. Biodiversity of Australian desert lizards. In C. I. Peng and C. H. Chou (eds.) Biodiversity and Terrestrial Ecosystems. Institute of Botany, Academica Sinica, Monograph Series No. 14, pp. 259-281. Download pdf

Pianka, E. R. 1995. Evolution of body size: Varanid lizards as a model system. American Naturalist 146: 398-414. Download pdf.

Pianka, E. R. 1995. Desert lizard ecology: Panglobal comparisons. Hardun. Journal of the Israel Herpetological Information Center, No. 6: 25-38. (in hebrew).

Pianka, E. R. 1996. Long-term changes in lizard assemblages in the Great Victoria Desert: Dynamic habitat mosaics in response to wildfires. Chapter 8 (pp. 191-215) in M. L. Cody and J. A. Smallwood (eds.) Long-term Studies of Vertebrate Communities. Academic Press. Download pdf

Pianka, G. A., E. R. Pianka, and G. G. Thompson. 1996. Egg laying by thorny devils (Moloch horridus) under natural conditions in the Great Victoria desert. J. Roy. Soc. Western Australia 79: 195-198. Download pdf

Perry, G. and E. R. Pianka. 1997. Animal foraging: past, present and future. Trends in Ecology and Evolution 12: 360-364. Download pdf

Couper, P. J. and E. R. Pianka. 1997. Synonymy of Ctenotus monticola Storr, 1981 and Ctenotus hypatia Ingram and Czechura, 1990. Memoirs of the Queensland Museum 42: 12.

Pianka, E. R. 1997. Australia's thorny devil. Reptiles 5 (11): 14-23. Download pdf

Pianka, E. R., D. M. Hillis, D. C. Cannatella, M. J. Ryan, and J. J. Wiens. 1998. Teaching Herpetology. Herpetologica 54 (Supplement): S3-S5. Download pdf

Pianka, E. R. and W. L. Hodges . 1998. Horned lizards. Reptiles 6 (6): 48-63. Translated into Finnish in 2003, published as Sarvikonnaliskot -- Phrynosoma in Herpetomania 12: 5-19.

Pianka, G. A., E. R. Pianka, and G. G. Thompson. 1998. Natural history of thorny devils Moloch horridus (Lacertilia: Agamidae) in the Great Victoria desert. J. Royal Society of Western Australia 81: 183- 190. Download pdf

Thompson, G. G., E. R. Pianka, and M. de Boer. 1999. Body temperatures of an arboreal monitor lizard, Varanus tristis (Squamata: Varanidae) during the breeding season. Amphibia- Reptilia 20: 82-88. Download pdf

Thompson, G. G. and E. R. Pianka. 1999. Reproductive ecology of the black-headed goanna Varanus tristis (Squamata: Varanidae). Journal of the Royal Society of Western Australia 82: 27-31. Download pdf

Thompson, G. G., M. de Boer, and E. R. Pianka. 1999. Activity areas and daily movements of an arboreal monitor lizard, Varanus tristis (Squamata: Varanidae) during the breeding season. Australian Journal of Ecology 24: 117-122. Download pdf

Pearson, D. J., D. R. King, and E. R. Pianka. 1999. New localities for the western pygmy possum, Cercartetus concinnus. Western Australian Naturalist 22: 205-207.

Haydon, D. T. and E. R. Pianka. 1999. Metapopulation theory, landscape models, and species diversity. EcoScience 6: 316-328. Abstract, Download pdf

Haydon, D. T., J. K. Friar, and E. R. Pianka. 2000. Fire Driven Dynamic Habitat Mosaics in the Great Victoria Desert I: Fire Geometry. Landscape Ecology 15: 373-381. Abstract, Download pdf

Haydon, D. T., J. K. Friar, and E. R. Pianka. 2000. Fire Driven Dynamic Habitat Mosaics in the Great Victoria Desert II: A spatial and temporal landscape model. Landscape Ecology 15: 407-423. Abstract, Download pdf

Farlow, J. O. and E. R . Pianka. 2000. Body form and trackway pattern in Australian desert monitors (Squamata: Varanidae): Comparing zoological and ichnological diversity. Palaios 15: 235-247. Abstract, Download pdf

Huey, R. B., E. R. Pianka, and L. J. Vitt. 2001. How often do lizards "run on empty?" Ecology 82: 1-7. Download pdf

Pianka, E. R. 2001. The wonderful lizards of Oz. Fauna 2: 8-17.

Thompson, G. G. and E. R. Pianka. 2001. Allometry of Clutch and Neonate Sizes in Monitor Lizards (Varanidae: Varanus). Copeia 2001: 443-458. Download pdf

Winemiller, K. O., E. R. Pianka, L. J. Vitt, and A. Joern. 2001. Food web laws or niche theory? Six independent empirical tests. American Naturalist 158: 193- 199. Download pdf

Pianka, E. R. 2001. The role of phylogenetics in evolutionary ecology. Pp. 1-20 in Herpetologia Candiana. P. Lymberakis, E. Valakos, P. Pafilis, and M. Mylonas (eds), Natural History Museum of Crete, Societas Europaea Herpetologica, Irakleio Crete 2001. Download pdf

Pianka, E. R. 2001. Evolutionary Stable Strategies. pp. 669-671 in Encyclopedia of Genetics, Academic Press. Download pdf

Pianka, E. R. 2002. Species Diversity. In The Oxford Encyclopedia of Evolution.

Pianka, E. R. 2002. Adaptation. In a 4 volume Encyclopedia, titled "Biology for Students Vol. II. Evolution." Reference Division, Macmillan Publishing. Link to read. Download pdf

Pianka, E. R. 2002. Convergent Evolution. In a 4 volume Encyclopedia, titled "Biology for Students Vol. II. Evolution." Reference Division, Macmillan Publishing. Link to read.

Pianka, E. R. 2002. Natural Selection. In a 4 volume Encyclopedia, titled "Biology for Students Vol. II. Evolution." Reference Division, Macmillan Publishing. Link to read.

Pianka, E. R. 2002. Forward. Pages xi-xiii in J. B. Murphy, C. Ciofi, Colomba de La Panouse and T. Walsh (eds.) Komodo dragons: Biology and conservation. Smithsonian Institute Press.

King, D. R., E. R. Pianka, and B. Green. 2002. Biology, ecology, and evolution. Chapter 2 (pages 23-41) in J. B. Murphy, C. Ciofi, Colomba de La Panouse and T. Walsh (eds.) Komodo dragons: Biology and conservation. Smithsonian Institute Press.

Pianka, E. R. 2002. A general review of zoological trends during the 20th century. A. Legakis, S. Sfenthourakis, R. Polymeni, and M. Thessalou-Legaki, eds. Proc. 18th International Congress of Zoology, pp.3-13. Opening address at the 18th International Congress of Zoology, Athens 2001. Abstract. Download pdf

Vitt, L. J., E. R. Pianka, W. E. Cooper, and K. Schwenk. 2003. History and the global ecology of squamate reptiles. American Naturalist 162: 44-60. Download pdf

Pianka, E. R. 2003. Squamata. In Grzimek's Animal Life Encyclopedia. pp. 195-208.

Pianka, E. R. 2003. Agamidae. In Grzimek's Animal Life Encyclopedia. pp. 209-222.

Pianka, E. R. 2003. Dibamidae. In Grzimek's Animal Life Encyclopedia. pp. 271-272.

Pianka, E. R. 2003. Scincidae. In Grzimek's Animal Life Encyclopedia. pp. 327-338.

Pianka, E. R. 2003. Varanidae. In Grzimek's Animal Life Encyclopedia. pp. 359-368.

Thompson, G. G., P. C. Withers, E. R. Pianka, and S. A. Thompson. 2003. Assessing biodiversity with species accumulation curves: Inventories of small reptiles by pit-trapping in Western Australia. Austral Ecology 28: 361-383. Download pdf

Thompson, G. G., S. A. Thompson, P. C. Withers and E. R. Pianka. 2003 Diversity and abundance of pit-trapped reptiles of arid and mesic habitats in Australia: Biodiversity for Environmental Impact Assessments. Pacific Conservation Biology 9: 120-135. Download pdf

Pianka, E. R. 2003. Australian Desert Varanids. Reptilia 30: 20-26. Translated in Europe into German and Spanish.

Pianka, E. R. and W. L. Hodges . 2003. Sarvikonnaliskot -- Phrynosoma. Herpetomania 12: 5-19 (in Finnish).

Sweet, S. S. and E. R. Pianka. 2003. The lizard kings. Natural History 112(9): 40- 45. Download pdf

Farlow, J. O. and E. R . Pianka. 2003. Body size overlap, habitat partitioning, and living-space requirements of terrestrial vertebrate predators: implications for large-theropod paleoecology. Historical Biology 16: 21-40. Download pdf

Jennings, W. B., E. R. Pianka, and S. Donnellan. 2003. Systematics of the lizard family Pygopodidae with implications for the diversification of Australian temperate biotas. Systematic Biology 52: 757-780. Download pdf

Vitt, L. J. and E. R. Pianka. 2003. Historical patterns in lizard ecology: what teiids can tell us about lacertids. In V. Perez-Mellado (ed.) Lacertids of the Mediterranean Basin. pp. 139-156. Download pdf

Pianka, E. R. and D. R. King. 2004. Introduction. Chapter 1 (pp. 3-9) in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press. Read selected pages on line

Molnar, R. E. and E. R. Pianka, E. R. 2004. Biogeography and Phylogeny of Varanoids. Chapter 3 (pp. 68-76) in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Jennings, W. B. and E. R. Pianka, E. R. 2004. Tempo and Timing of the Australian Varanus Radiation. Chapter 4 (pp. 77-87) in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Varanus bengalensis. Pages 156-160 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Varanus olivaceus. Pages 220-224 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Varanus brevicauda. Pages 312-317 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Varanus eremius. Pages 328-334 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Varanus semiremex. Pages 462-465 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Varanus tristis. Pages 477-487 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Lanthanotus borneensis. Pages 535-538 in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. 2004. Evolution of Body Size and Reproductive Tactics. Chapter 9 (pp. 549-555) in Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press.

Pianka, E. R. and S. S. Sweet. 2005. Integrative biology of sticky feet in geckos. BioEssays 27: 647-652. Download pdf

Vitt, L. J. and E. R. Pianka. 2005. Deep history impacts present day ecology and biodiversity. Proc. Nat. Acad Sci. 102:7877-7881. Download pdf. Download pdf of commentary by Harry Greene

Pianka, E. R. 2005. Zur Okologie und Naturgeschichte der Zwergbartagame (Pogona minor) in der Great Victoria Desert (Australien). Draco: 63-66. Download pdf in English.

Pianka, E. R. and H. S. Horn. 2005. Ecology's Legacy from Robert MacArthur. Chapter 11 (pp. 213-232) in K. Cuddington and B. Biesner, eds. "Ecological Paradigms Lost: Routes of Theory Change." Elsevier Academic Press.

Hibbitts, T. J., E. R. Pianka, R. B. Huey, and M. J. Whiting. 2005. Ecology of the Common Barking Gecko (Ptenopus garrulus) in southern Africa. J. Herpetology 39: 509-515. Download pdf

Vitt, L. J. and E. R. Pianka. 2006. The Scaly Ones. Natural History. 115 (6): 28-35. Download pdf

Pianka, E. R. 2007. Molokki -- Piikikas paholainen. Herpetomania 16: 5-11 (in Finnish).

Huey, R. B. and E. R. Pianka. 2007. On Widely Foraging for Kalahari Lizards. Preface, pp. 1-10 in S. M. Reilly, L. D. McBrayer, and D. B. Miles, eds. "Lizard Ecology: The Evolutionary Consequences of Foraging Mode." Cambridge University Press. Read on line Download pdf

Vitt, L. J. and E. R. Pianka. 2007. Feeding Ecology in the Natural World. Chapter 5, pp. 141-172 in S. M. Reilly, L. D. McBrayer, and D. B.Miles, eds. "Lizard Ecology: The Evolutionary Consequences of Foraging Mode." Cambridge University Press.

Huey, R. B. and E. R. Pianka. 2007. Lizard thermal biology: do genders differ? Amer. Natur.170: 473-478. Download pdf

Sweet, S. S. and E. R. Pianka. 2007. Monitors, mammals and Wallace's Line. Third multidisciplinary world conference on monitor lizards, Alexander Koenig Museum, Bonn, Germany. Mertensiella 16: 79-99. Download pdf

King, D. R. and E. R. Pianka. 2007. Ecology of the Pygmy Monitor Varanus brevicauda in Western Australia. Third multidisciplinary world conference on monitor lizards, Alexander Koenig Museum, Bonn, Germany. Mertensiella 16: 304-311. Download pdf

Pianka, E. R. 2007. An update on the Ecology of the Pygmy Monitor Varanus eremius in Western Australia. Third multidisciplinary world conference on monitor lizards, Alexander Koenig Museum, Bonn, Germany. Mertensiella 16: 346-352. Download pdf

Pianka, E. R. 2008. Autecology. In Jorgensen, S. E., ed. Encyclopedia of Ecology. Elsevier.

Pianka, E. R. 2008. Evolutionary Ecology. In Jorgensen, S. E., ed. Encyclopedia of Ecology. Elsevier.

Pianka, E. R. 2008. Optimal Foraging. In Jorgensen, S. E., ed. Encyclopedia of Ecology. Elsevier.

Pianka, E. R. 2008. Reproductive Tactics. In Jorgensen, S. E., ed. Encyclopedia of Ecology. Elsevier.

Pianka, E. R. 2008. r and K selection. In Jorgensen, S. E., ed. Encyclopedia of Ecology. Elsevier.

Pianka, E. R. 2008. Synecology. In Jorgensen, S. E., ed. Encyclopedia of Ecology. Elsevier.

Goodyear, S. E. and E. R Pianka. 2008. Sympatric Ecology of Five Species of Fossorial Snakes (Elapidae) in Western Australia. J. Herpetology 42: 279-285. Download pdf

Costa, G. C., L. J. Vitt, E. R. Pianka, D. O. Mesquita and G. R. Colli. 2008. Optimal foraging constrains macroecological patterns: Body size and dietary niche breadth in lizards. Global Ecology and Biogeography 17: 670-677. Download pdf


Harp, C. A., S. E. Goodyear, and E. R. Pianka. 2010. Ctenotus helenae (Helen's Comb-Eared Skink) Hermaphroditism. Herpetological Review 41: 353-354. Download pdf



Pianka, E. R. 2010. Notes on the ecology of four species of pygopodid lizards in the Great Victoria Desert. Western Australian Naturalist 27: 175-181. Download pdf

Pianka, E. R. 2011. Scientific Methods and Human Knowledge.
Pp. 1-7 in M. Hutchins, V. Geist, and E. R. Pianka, eds. Grzimek's Animal Life Encyclopedia, Evolution. Gale Publishing Group.

Pianka, E. R. 2011. Convergent evolution and ecological equivalence. Pp. 235-242 in M. Hutchins, V. Geist, and E. R. Pianka, eds. Grzimek's Animal Life Encyclopedia, Evolution. Gale Publishing Group. Download pdf

Pianka, E. R. 2011. Notes on the natural history of
the tiny skink Menetia greyi. Western Australian
Naturalist 28: 12-17. Download pdf


Pianka, E. R. and C. A. Harp. 2011. Notes on the natural history of Buchanan's snake-eyed skink Cryptoblepharus buchananii in arid Western Australia. Western Australian Naturalist 28: 43-49. Download pdf

Pianka, E. R. 2011. Notes on the ecology of some uncommon skinks in the Great Victoria Desert. Western Australian Naturalist 28: 50-60. Download pdf


Positions in the first two principal components of dietary niche space for 4 abundant species of Ctenotus skinks from 3 sites for each of 5 time intervals (those at the B-area are joined by lines forming polygons).

The first 2 components reduce variation by 60% (PC3 contributes a further 13%, not shown). Positions of five major food items, Hemiptera (true bugs), Orthoptera (grasshoppers), Aranae (spiders), insect larvae, and Isoptera (termites) are shown. PC1 loads primarily on a Hemiptera-Isoptera axis whereas PC2 loads on a Hemiptera-Orthoptera-Aranae axis. Ctenotus piankai eats more hemiptera than other species and Ctenotus calurus consumes more termites. Ctenotus quattuordecimlineatus eats more grasshoppers and spiders, and Ctenotus pantherinus occupies the central part of this dietary niche space. Diets shift in time but each species occupies its own distinct zone. Each point can be considered an estimate of the realized food niche whereas the area filled by all data points for each species represents its fundamental food niche. Overlap between species occurs but is limited (Goodyear and Pianka 2011).


Goodyear, S. E. and E. R. Pianka. 2011. Spatial and temporal variation in diets of sympatric lizards (genus Ctenotus) in the Great Victoria Desert, Western Australia. J. Herpetology 45: 265-271. Read On Line. Download pdf

Pianka, E. R. and S. E. Goodyear. 2012. Lizard responses to wildfire in arid interior Australia: Long-term experimental data and commonalities with other studies. Austral Ecology 37: 1-11. Download pdf

Pianka, E. R. 2012. Can humans share spaceship earth? (“Point of View”) Amphibian and Reptile Conservation 6(1): 1-24(e49). Download pdf (2.8 megs).

Böhm M. et al. (with co­authors including E. R. Pianka). 2013. The conservation status of the world’s reptiles. Biological Conservation 157: 372-385. Download pdf


Pianka, E. R. 2013. Notes on the ecology and natural
history of two uncommon arboreal agamid lizards
Diporiphora paraconvergens and Lophognathus
longirostris
in the Great Victoria desert of Western
Australia. Western Australian Naturalist 29: 77-84.
Download pdf


Pianka, E. R. 2013. Notes on the ecology and natural history of two uncommon terrestrial agamid lizards Ctenophorus clayi and C. fordi in the Great Victoria desert of Western Australia. Western Australian Naturalist 29: 85-93. Download pdf

Pianka, E. R. 2013. Notes on the natural history of the rarely recorded agamid lizard Caimanops amphiboluroides in Western Australia. Western Australian Naturalist 29: 99-102. Download pdf

Pianka, E. R. 2013. Notes on the ecology and natural history of the rarely recorded gekkonid lizard Heteronotia binoei in the Great Victoria desert of Western Australia. Western Australian Naturalist 29: 120-125. Download pdf

Pianka, E. R. 2014. Rarity in Australian Desert Lizards. Austral Ecology 39: 214-224. pdf

Pianka, E. R. 2014. Notes on a collection of lizards from the Eucla sand dunes in Western Australia. Western Australian Naturalist 30: 155-161..

Pianka, E. R. 2014. Notes on the ecology and natural history of Ctenophorus reticulatus (Agamidae) in Western Australia. Western Australian Naturalist 30: 222-225..

Pianka, E. R. 2014. Notes on the ecology and natural history of Ctenophorus caudicinctus (Agamidae) in Western Australia. Western Australian Naturalist 30: 226-230..

Pianka, E. R. 2014. Notes on the ecology and natural history of two species of Egernia (Scincidae) in Western Australia. Western Australian Naturalist 30: 231-236..




Books by Eric R. Pianka



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